Description: The Himalaya and the Tibetan Plateau are uplifted by the ongoing northward underthrusting of the Indian continental lithosphere below Tibet resulting in lithospheric stacking. The layered structure of the Tibetan upper mantle is imaged by seismic methods, most detailed with the receiver function method. Tibet is considered as a place where the development of a future craton is currently under way. Here we study the upper mantle from Germany to northern Sweden with seismic S receiver functions and compare the structure below Scandinavia with that below Tibet. Below Proterozoic Scandinavia, we found two low velocity zones on top of each other, separated by a high velocity zone. The top of the upper low velocity zone at about 100km depth extends from Germany to Archaean northern Sweden. It agrees with the lithosphere-asthenosphere boundary (LAB) below Germany and Denmark. Below Sweden it is known as the 8°discontinuity, or as a mid-lithospheric discontinuity (MLD), similar to observations in North America. Seismic tomography places the LAB near 200km in Scandinavia, which is close to the top of our deeper low velocity zone. We also observed the bottom of the asthenosphere (the Lehmann discontinuity) deepening from 180km in Germany to 260km below Sweden. Remnants of old subduction in the upper about 100km below Scandinavia and Finland are known from controlled source seismic experiments and local earthquake studies. Recent tomographic studies indicate delamination of the lithosphere below southern Scandinavia and northern Germany. We are suggesting that the large scale layered structure in the Scandinavian upper mantle may be caused by processes similar to the ongoing lithospheric stacking in Tibet.

Description: Teleost fish are among the most ancient vertebrates possessing an adaptive immune system with B and T lymphocytes that produce memory responses to pathogens. Most bony fish, however, have only 2 types of B lymphocytes, in contrast to the 4 types available to mammals. To better understand the evolution of adaptive immunity, we generated transgenic zebrafish in which the major immunoglobulin M (IgM + ) B-cell subset expresses green fluorescence protein (GFP) ( IgM1:eGFP ). We discovered that the earliest IgM + B cells appear between the dorsal aorta and posterior cardinal vein and also in the kidney around 20 days postfertilization. We also examined B-cell ontogeny in adult IgM1:eGFP;rag2:DsRed animals, where we defined pro-B, pre-B, and immature/mature B cells in the adult kidney. Sites of B-cell development that shift between the embryo and adult have previously been described in birds and mammals. Our results suggest that this developmental shift occurs in all jawed vertebrates. Finally, we used IgM1:eGFP and cd45DsRed;blimp1:eGFP zebrafish to characterize plasma B cells and investigate B-cell function. The IgM1:eGFP reporter fish are the first nonmammalian B-cell reporter animals to be described. They will be important for further investigation of immune cell evolution and development and host-pathogen interactions in zebrafish.

Description: Plate tectonic reconstructions are usually constrained by the correlation of lineaments of surface geology and crustal structures. This procedure is, however, largely dependent on and complicated by assumptions on crustal structure and thinning and the identification of the continent-ocean transition. We identify two geophysically and geometrically similar upper mantle structures in the North Atlantic and suggest that these represent remnants of the same Caledonian collision event. The identification of this structural lineament provides a sub-crustal piercing point and hence a novel opportunity to tie plate tectonic reconstructions. Further, this structure coincides with the location of some major tectonic events of the North Atlantic post-orogenic evolution such as the occurrence of the Iceland Melt Anomaly and the separation of the Jan Mayen microcontinent. We suggest that this inherited orogenic structure played a major role in the control of North Atlantic tectonic processes.

Description: The postorogenic collapse of the early Paleozoic Caledonian orogeny is well documented; however, several different plate tectonic models exist for the convergent phase involving closure of the Iapetus Ocean and the collision of Laurentia and Baltica. Receiver function analysis of 11 broadband seismometers along a 270 km transect in the East Greenland Caledonides reveals the existence of an east-dipping high velocity slab. Numerical modeling demonstrates that relict subducted and eclogitized crust is a plausible explanation. Thus, eastward subduction preceded subsequent west-dipping subduction during the formation of the East Greenland and Scandinavian Caledonides. This is a key constraint for understanding the Caledonian and continental margin evolution in the North Atlantic realm.

Description: Ellesmere Island in Arctic Canada displays a complex geological evolution. The region was affected by two distinct orogenies, the Palaeozoic Ellesmerian orogeny (the Caledonian equivalent in Arctic Canada and Northern Greenland) and the Palaeogene Eurekan orogeny, related to the opening of Baffin Bay and the consequent convergence of the Greenland plate. The details of this complex evolution and the present-day deep structure are poorly constrained in this remote area and deep geophysical data are sparse. Receiver function analysis of seven temporary broad-band seismometers of the Ellesmere Island Lithosphere Experiment complemented by two permanent stations provides important data on the crustal velocity structure of Ellesmere Island. The crustal expression of the northernmost tectonic block of Ellesmere Island (~82°–83°N), Pearya, which was accreted during the Ellesmerian orogeny, is similar to that at the southernmost part, which is part of the Precambrian Laurentian (North America-Greenland) craton. Both segments have thick crystalline crust (~35–36 km) and comparable velocity–depth profiles. In contrast, crustal thickness in central Ellesmere Island decreases from ~24–30 km in the Eurekan fold and thrust belt (~79.7°–80.6°N) to ~16–20 km in the Hazen Stable Block (HSB; ~80.6°–81.4°N) and is covered by a thick succession of metasediments. A deep crustal root (~48 km) at ~79.6°N is interpreted as cratonic crust flexed beneath the Eurekan fold and thrust belt. The Carboniferous to Palaeogene sedimentary succession of the Sverdrup Basin is inferred to be up to 1–4 km thick, comparable to geologically-based estimates, near the western margin of the HSB.

Description: This study presents P - and S -wave velocity variations for the upper mantle in southern Scandinavia and northern Germany based on teleseismic traveltime tomography. Tectonically, this region includes the entire northern part of the prominent Tornquist Zone which follows along the transition from old Precambrian shield units to the east to younger Phanerozoic deep sedimentary basins to the southwest. We combine data from several separate temporary arrays/profiles (276 stations) deployed over a period of about 15 yr and permanent networks (31 stations) covering the areas of Denmark, northern Germany, southern Sweden and southern Norway. By performing an integrated P - and S -traveltime analysis, we obtain the first high-resolution combined 3-D V P and V S models, including variations in the V P / V S ratio, for the whole of this region of study. Relative station mean traveltime residuals vary within ±1 s for P wave and ±2 s for S wave, with early arrivals in shield areas of southern Sweden and later arrivals in the Danish and North German Basins, as well as in most of southern Norway. In good accordance with previous, mainly P -velocity models, a marked upper-mantle velocity boundary (UMVB) is accurately delineated between shield areas (with high seismic mantle velocity) and basins (with lower velocity). It continues northwards into southern Norway near the Oslo Graben area and further north across the Southern Scandes Mountains. This main boundary, extending to a depth of at least 300 km, is even more pronounced in our new S -velocity model, with velocity contrasts of up to ±2–3 per cent. It is also clearly reflected in the V P / V S ratio. Differences in this ratio of up to about ±2 per cent are observed across the boundary, with generally low values in shield areas to the east and relatively higher values in basin areas to the southwest and in most of southern Norway. Differences in the V P / V S ratio are believed to be a rather robust indicator of upper-mantle compositional differences. For the depth interval of about 100–300 km, thick, depleted, relatively cold shield lithosphere is indicated in southern Sweden, contrasting with more fertile, warm mantle asthenosphere beneath most of the basins in Denmark and northern Germany. Both compositional and temperature differences seem to play a significant role in explaining the UMVB between southern Norway and southern Sweden. In addition to the main regional upper-mantle velocity contrasts, a number of more local anomaly features are also outlined and discussed.

Description: The large-scale geological evolution of the North Atlantic Realm during the past 450 Myr is largely understood, but crucial elements remain uncertain. These involve the Caledonian orogeny, the formation of the North Atlantic and accompanying igneous activity, and the present-day high topography surrounding the North Atlantic. Teleseismic receiver function interpretation in the Central Fjord Region of East Greenland recently suggested the presence of a fossil Caledonian subduction complex, including a slab of eclogitised mafic crust and an overlying wedge of serpentinised mantle peridotite. Here we further investigate this topic using inverse receiver functions modelling. The obtained velocity models are tested with regard to their consistency with the regional gravity field and topography. We find that the obtained receiver function model is generally consistent with gravity and isostasy. The western part of the section, with topography of 〉1000 m, is clearly supported by the 40-km-thick crust. The eastern part requires additional buoyancy as provided by the hydrated mantle wedge. The geometry, velocities and densities are consistent with interpretation of the lithospheric structure as a fossil subduction zone complex. The spatial relations with Caledonian structures suggest a Caledonian origin. The results indicate that topography is isostatically compensated by density variations within the lithosphere, and that significant dynamic topography is not required at the present-day.

Description: The relative traveltime residuals of more than 20 000 arrival times of teleseismic P and S waves measured over a period of more than 10 yr in five separate temporary and two permanent seismic networks covering the Scandinavian (Scandes) Mountains and adjacent areas of the Baltic Shield are inverted to 3-D tomograms of P and S velocities and the V P / V S ratio. Resolution analysis documents that good 3-D resolution is available under the dense network south of 64° latitude (Southern Scandes Mountains), and patchier, but highly useful resolution is available further north, where station coverage is more uneven. A pronounced upper-mantle velocity boundary (UMVB) that transects the study region is defined. It runs from SE Norway (east of the Oslo Graben) across the mountains to the Norwegian coast near Trondheim (around the Møre–Trøndelag Fault Complex), after which it follows closely along the coast further north. Seismic velocities in the depth interval 100–300 km change significantly across the UMVB from low relative V P and even lower relative V S on the western side, to high relative V P and even higher relative V S to the east. This main velocity boundary therefore also separates relatively high V P / V S ratio to the west and relatively low V P / V S to the east. Under the Southern Scandes Mountains (most of southern Norway), we find low relative V P , even lower relative V S and hence high V P / V S ratios. These velocities are indicative of thinner lithosphere, higher temperature and less depletion and/or fluid content in a relatively shallow asthenosphere. At first sight, this might support the idea of a mantle buoyancy source for the high topography. Under the Northern Scandes Mountains, we find the opposite situation: high relative V P , even higher relative V S and hence low V P / V S ratios, consistent with thick, dry, depleted lithosphere, similar to that in most of the Baltic Shield area. This demonstrates significant differences in upper-mantle conditions between the Southern and Northern Scandes Mountains, and it shows that upper-mantle velocity anomalies are very poor predictors of topography in this region. An important deviation from this principal pattern is found near the topographic saddle between the Southern and Northern Scandes Mountains. Centred around 64°N, 14°E, a zone of lower S velocity and hence higher V P / V S ratio is detected in the depth interval between 100 and 300 km. This ‘Trøndelag–Jämtland mantle anomaly’ (TJMA) is still interpreted as part of relatively undisturbed lithosphere of shield affinity because of high relative P velocity, but the relatively low V P / V S ratios indicate lower depletion, possibly higher fluid content, and most likely lower viscosity relative to the adjacent shield units. We suggest that this mantle anomaly may have influenced the collapse of the Caledonian Mountains, and in particular guided the location and development of the Møre–Trøndelag Fault Complex. The TJMA is therefore likely to have played an important role in the development of the ‘two-dome architecture’ of the Scandes Mountains.