Description: A high-resolution calcareous nannofossil analysis of the Danian/Selandian boundary was conducted at Site 1262 (Walvis Ridge, South Atlantic) to pinpoint the lowest occurrence of fasciculiths and to unravel the evolutionary trends throughout nannofossil Zone NP4. Using quantitative analyses, numerous primary and secondary bioevents were identified, improving the biostratigraphic resolution of this interval. The main events recorded at Site 1262 were also identified at the Zumaia section Global Stratotype Section and Point (GSSP) of the base of the Selandian and at the Qreiya section (Egypt). The lowest occurrence of fasciculiths (represented by the LO of Gomphiolithus magnicordis and Gomphiolithus magnus) was observed in the middle part of Chron C27r, above the LO of Toweius pertusus and prior to the LO of the genus Sphenolithus. The synchroneity of the LO of fasciculiths was also verified at various latitudes, such as DSDP Site 384 (NW Atlantic), ODP Site 761B (Indian Ocean) and DSDP Site 577A (Pacific Ocean). The first and second diversification events (Steurbaut and Sztrákos, 2008, doi:10.1016/j.marmicro.2007.08.004), or radiation events (Bernaola et al., 2009, doi:10.1344/105.000000272), of fasciculiths have been thoroughly discussed and well characterized by a succession of events. The occurrence of the Latest Danian Event (LDE) and several paleoenvironmental changes recognized during this time interval, coupled with an ecological competition with Sphenolithus, appear to be the probable causes of the First and Second Radiations and the fasciculith barren interval between them. The occurrence of new morphostructures and taxa suggests evolutionary trends and a strict link between morphological evolution and paleoclimate.