Description: Highlights • Hikurangi Plateau crust at the southern Hikurangi margin is ≈12 km thick. • A fast-upper mantle resides at a depth of ≈50 km beneath Hikurangi trough. • Crustal and upper mantle P-wave-speeds reduced by ≈10% beneath the Hikurangi trough. • Hikurangi Plateau is hydrated to depths of ≈50 km beneath the Hikurangi trough. Abstract Controlled-source seismic studies at most subduction zones show that bending of the subducting plate results in reduced seismic wave-speeds in the crust and upper mantle near the trench. Similar studies also have found unusually high P-wave-speeds (Vp) in the upper mantle under oceanic plateaus. Onshore-offshore seismic profiling at the southern Hikurangi margin, where the ≈120 Ma old oceanic Hikurangi Plateau is subducting, indicates that a fast (Vp≈8.7±0.2 km/s) upper mantle layer lies beneath a ≈25 km thick mantle layer with more regular wave-speeds (Vp≈8.0±0.2 km/s) under the Hikurangi trough. This is consistent with previous findings of upper mantle Vp≈8.7-9.0 km/s in the margin-parallel direction under the North Island (≈100 km northwest of the deformation front) at depths ≈8-10 km below the Moho. Our profiles are margin-perpendicular, thus we show that the upper mantle lid of the subducting Pacific Plate is characterized by unusually high P-wave-speeds along all azimuths. We find an area of lowered Vp in the ≈12±1 km thick Hikurangi Plateau crust beneath the trough. This drop in Vp is ≈10%, and a similar drop in Vp is deduced to depths of 25±2 km into the upper mantle. We interpret that the increase in thickness of the regular mantle beneath the trough results from the formation of a low-velocity zone in the faster upper mantle layer; this zone formed from serpentinisation by hydration through bending-induced normal faults and/or due to crack porosity introduced by thermal cracking, further enhanced by bending-related faulting. Thus the “regular mantle” (Vp≈8 km/s) is not in fact regular, but rather the high-speed mantle has mechanically bent, fractured, and altered. The absolute depth of fast mantle Vp under the Hikurangi trough is around 50 km. The onset of the lower band of seismicity of the double seismic zone and high upper mantle Vp under the North Island is observed at similar depths. This is consistent with the hypothesis that the lower band of earthquakes in a double seismic zone is due to antigorite dehydration processes, a hydrous mineral formed in the low velocity zone in the upper mantle beneath the trough. Our study on the Hikurangi margin is different, as the subducting plate here contains a ≈120 Ma old oceanic plateau with a ≈12 km thick crust, but the results are similar to other subduction margins where regular oceanic crust is subducting.