Description: Côte d’Ivoire produces 40 % of the world supply of cocoa but much of the plantation area is aging and declining in productivity, while opportunities for land expansion into new forest land are quickly disappearing. Rejuvenation strategies for cocoa presently coalesce either around improved varieties and greater use of agro-chemical inputs in full sun systems or eco-certification that requires trees to be integrated with cocoa. Here, we explore the possibility of uniting these approaches through building on current farmer practice of incorporating trees in their cocoa fields to improve cocoa productivity and diversify their livelihoods. We interviewed 355 farmers about trees integrated in their cocoa fields across four locations in the South-West of Côte d’Ivoire in 2012, stratified by whether or not farmers were eco-certified. Despite the massive deforestation, a rich diversity of trees was found in cocoa fields and an overwhelming majority of farmers (95 %) wanted more trees and/or more tree species, regardless of their certification status or ethnic origin. There was a consensus that most trees were compatible with cocoa, but farmers also traded off negative impacts of some species against their productive contribution to their livelihood. Farmers valued tree diversity on their cocoa plots and provided detailed information on how 32 tree species interacted with cocoa in terms of soil moisture retention, soil fertility improvement and pest and disease interactions but also had key gaps in knowledge about alternative hosts of mirids and mistletoe. The majority of farmers were not aware of the certification requirements for tree species and shade cover but a much higher proportion of certified farmers (76 %) had received information about shade trees than non-certified farmers (15 %), although advice only related to eight tree species. Scope for building on local knowledge and practice to sustainably increase cocoa productivity through promoting tree diversity while enhancing other ecosystem service provision was identified and the next steps required to realize this set out.

Description: The benefits of tree-based intercropping (TBI) compared to conventional agro-ecosystems in North America could include climate change mitigation and adaptation, although enhancing resilience to climate change through increasing soil diversity remains poorly explored. Diversity of soil microarthropods supports a series of ecological services that may be altered by soil desiccation due to climate change. Here we study the effect of red oak and hybrid poplar TBI on soil oribatid mite species assemblages associated to forage crops (mix of Timothy-grass and red clover). Abundance and species density of oribatids were affected by treatment, depth and the interaction of both variables. Abundance of oribatid mites was significantly lower in the oak TBI, showing a homogeneous vertical distribution in opposition to a decreasing with depth distribution under poplar TBI and conventional crops. Species density was significantly higher in the conventional crop, showing again significant differences in depth that were not present in both TBI treatments. Distance to tree did not affect mite abundance nor species density. TBI increased oribatid richness (obtained by sample-based rarefaction and extrapolation) only in the presence of oaks. The distribution of oribatids was strongly associated to tree fine root biomass and stress the importance of underground organic resources for the oribatid fauna and their ecological functions. If increasing drought associated with climate change desiccates superficial levels of agro-ecosystem soils, deeper sources of organic resources, such as tree roots, should become crucial in the maintenance of diverse microarthropod communities.

Description: In this study we analysed: (1) the biodiversity conservation capacity of Agroforestry Systems (AFS) in temperate highlands of the Tehuacán–Cuicatlán Valley, Central Mexico, (2) human cultural motives and actions for conserving such diversity and (3) problems endangering that capacity. We evaluated the richness and diversity of perennial plant species maintained in AFS through vegetation sampling of 14 agricultural plots and compared their composition with that of natural forests (14 plots of 500 m 2 each). We examined the situations among communities of Náhuatl, Ixcatec and Cuicatec people, documenting through interviews the management practices of plant species and the whole system, reasons why people maintain vegetation cover within AFS, and factors influencing changes in decisions favouring agriculture intensification. In the AFS studied we recorded a total of 79 species of trees and shrubs, 86 % of them being native species and representing 43 % of all species of trees and shrubs recorded in the sampling of the natural forests the AFS derive from. People leave standing on average a total of 40 individual trees and shrubs per agricultural plot. Reasons for leave plant species standing were more frequently associated with their use as fruit trees, firewood, shade, beauty, respect to nature and other environmental benefits. Water availability for irrigation, land tenure, and dependence on agriculture and forest for peasant's subsistence were main decision factors influencing AFS variation in their composition. AFS in temperate zones are important reservoirs of biodiversity and biocultural heritage and should be keystones for conservation policies in the Tehuacán–Cuicatlán Valley.

Description: Pastures are a major soil cover in Central Brazil, especially in rich soils previously occupied by dry forests. We simulated a scenario in which the wooden fences in Paranã Valley are replaced by live fences and isolated trees are left in the pasture fields, and we verified changing in tree cover by adding trees and avoiding logging for wooden fences. The simulation involved the analysis of a 20-year historic series of LANDSAT satellite images to determine the average time of pasture renewal. The average amount of wooden fences produced per hectare of local forest was estimated based on the literature and field data. The high spatial-resolution satellite images available in the Google Earth™ program were analyzed to estimate the total length of the fences and the average and maximum number of isolated trees per hectare found in the pastures of the region. The results showed that pasture renewal happens every 8.1 years. It is possible to produce an average of 1,472 stakes per hectare of forest. In the study area, we estimated the existence of an average of 842 km of wooden fences and 3.9 isolated trees per hectare of pasture (maximum = 48 isolated trees). The results of the simulation showed that the adoption of live fences can increase the crown coverage up to 7.5 % or even up to 14.3 % if all of the pasturelands are managed to have live fences and farmers begin to adopt cover-development practices, such as keeping an average of 48 isolated trees per hectare of pasture.

Description: This study evaluates weed dynamics during the first 4 years (2008–2011) of the implementation of an agroforestry system in the municipality of Tomé-açu in the state of Pará, Brazil. The study was conducted in a degraded pasture of Brachiaria humidicola , and treatments (T) included mixed plantations of oil palm with short-cycle leguminous species (T1) with the inclusion of manioc in T2 and forest species and palm trees in T3. In 2008, a floristic survey was conducted in 12 plots of 2 × 2 m per treatment. In 2009, herbicide was applied to one half of the area in each treatment. A total of 19,367 individuals of height 〈1.5 m were surveyed, of which 51 % were B. humidicola and 21 % were leguminous species that had been planted. Overall, an increase in plant density was observed between 2009 and 2010, followed by a decrease over the following year in areas without herbicide and a tendency for high densities to persist in areas with herbicide. Species richness increased over the study period in all areas (with or without herbicide), which might be related to habitat availability resulting from the reduced density of B. humidicola due to the effects of the herbicide, the leguminous species planted and the development of the agroforestry system. The association of short-cycle leguminous plants with afforestation appears to be an effective method for controlling weeds in the absence of herbicides.

Description: Even though agronomists have considered the spatial root distribution of plants to be important for interspecific interactions in agricultural intercropping, few experimental studies have quantified patterns of root distribution and their impacts on interspecific interactions in agroforestry systems. A field experiment was conducted to investigate the relationship between root distribution and interspecific interactions between intercropped jujube tree ( Zizyphus jujuba Mill.) and wheat ( Triticum aestivum Linn.) in Hetian, south Xinjiang province, northwest China. Roots were sampled by auger in 2-, 4- and 6-year-old jujube tree/wheat intercropping and in sole wheat and 2-, 4- and 6-year-old sole jujube down to 100 cm depth in the soil profile. The roots of both intercropped wheat and jujube had less root length density (RLD) at all soil depths than those of sole wheat and jujube trees. The RLD of 6-year-old jujube intercropped with wheat at different soil depths was influenced by intercropping to a smaller extent than in other jujube/wheat intercropping combinations. 6-year-old jujube exhibited a stronger negative effect on the productivity of wheat than did 2- or 4-year-old jujube and there was less effect on productivity of jujube in the 6-year-old system than in the 2- or 4-year-old jujube trees grown in monoculture. These findings may partly explain the interspecific competition effects in jujube tree/wheat agroforestry systems.

Description: The effect of tree canopy on the growth, productivity and forage quality of Megathyrsus maximus and changes in soil properties were evaluated over three seasonal periods. Four adjacent plots (15 m × 17 m each) in a tropical secondary deciduous forest having 12 years of growth and dominated by Gliricidia sepium were randomly assigned to two treatments: removal of trees (SCA) in two of the plots and leaving trees intact (COA) in the other two. In all plots, M . maximus was planted with 50 cm spacing among plants. Tree removal significantly increased the incident photosynthetically active radiation (PAR, P 〈 0.001) and grass size (12.5 % in height, P 〈 0.01, and 16.5 % in clump diameter, P 〈 0.05), but did not significantly affect any other variable. Season significantly affected grass height (P 〈 0.003), tiller number (P 〈 0.001), clump diameter (P 〈 0.001), net CO 2 assimilation rate (P 〈 0.001), forage biomass production (P 〈 0.003), and acid detergent fiber content (P = 0.033). Primary soil changes after 1 year of establishment of the grass were the decline by 3 % in organic carbon (P = 0.03), and qualitative changes in soil structure, regardless of tree presence. Results are consistent with the ability of M. maximus to tolerate shade. We conclude that under the conditions of the study there was no evidence for a negative effect of tree canopy on M. maximus mediated by a reduction in PAR.

Description: Twenty weeks forage re-growth from 20 multipurpose trees and shrubs (MPTS) species ( Albizia ferruginea , Albizia gummifera , Albizia niopoides , Berlinia grandiflora , Bauhimia monandra , Dialium guineense , Dalbergia sissoo , Enterolobium cyclocarpum , Leucaena leucocephala , Lonchocarpus sericeus , Milletia griffoneanus , Milletia thoningii , Napoliana imperialis , Parkia bicolor , Pterocarpus santalinoides , Senna spectabilis , Treculia africana , Terminalia superba , Tetrapluera tetraptera and Xylia xylocarpa ) were investigated for their feed value. The MPTS were grown on an ultisol in southeastern Nigeria and were subjected to proximate analyses, nylon bag degradability studies and cluster analysis. Among species, crude protein (CP) ranged from 104 g kg −1 DM in P. bicolor to 205 g kg −1 DM in A. gummifera , neutral detergent fibre (NDF) ranged from 498 g kg −1 DM in L. leucocephala to 771 g kg −1 DM in L. sericeus , acid detergent fibre (ADF) ranged from 303 g kg −1 DM in T. superba to 661 g kg −1 DM in L. sericeus , acid detergent lignin (ADL) ranged from 75 g kg −1 DM in T. superba to 305 g kg −1 DM in L. sericeus. Also, ADF-ash ranged from 0.78 g kg −1 DM in E. cyclocarpum to 15.58 g kg −1 DM in D. guineense. Except E. cyclocarpum and S. spectabilis , the remaining 18 MPTS studied had effective degradabilities (ED) less than 500 g kg −1 DM effective degradability. Eight MPTS ( A. ferruginea , A. niopoides , B. monandra , D. sissoo , E. cyclocarpum , L. leucocephala , S. spectabilis , T. superba ) were classified as high quality, another eight ( A. gummifera , B. grandiflora , L. sericeus , N. imperialis , P. bicolor , P. santalinoides , T. tetraptera and X. xylocarpa ) as medium quality while, the remaining four ( D. guineensis , M. griffoneanus , M. thoningii , T. africana ) fell into the low quality class based on their nutrient composition and ruminal DM degradability. The results indicate that most of the MPTS could be used as good quality feed for small ruminants in Nigeria.

Description: Despite of being structurally simpler and species-impoverished than natural forests, agroforestry plantations can act as a secondary habitat for native species and sustain some biodiversity. In particular, insectivores can provide important ecosystem services such as insectivory, indirectly benefiting plants through the reduction of herbivory and increasing productivity, by diminishing herbivores abundance. The lending of these services could occur in agroforestry systems, and in the same magnitude that in natural environments, however it has not yet been assessed whether the direct effect of insectivores over insects and indirect effect on plants differ between natural and agroforestry environments depending on possible modulating factors such as climatic region, type of insectivore, trophic group of the preyed arthropod and length of insectivores experimental exclusion. In this study, through a meta-analysis, it was assessed the provision of ecosystem services by insectivores in agroforestry systems compared to natural systems, contrasting it with the modulating factors mentioned. In general, insectivorous species reduced arthropod abundance and plant herbivory, and increased plant productivity. The magnitude and direction of these effects did not differ between natural and agroforestry systems, and neither did between different climatic regions, type of insectivore, preyed arthropod trophic groups nor experiment length. The effect of insectivores on productivity can vary based on the variable used to measure productivity. Our results evidence the provision of services by the insectivores present in plantations, independently of factors that could modulate its magnitude and direction. In this way, enhancing the existence of these important interactions within plantations could represent a win–win scenario.

Description: The Millwood (MW) cultivar of the honeylocust ( Gleditsia triacanthos L.) tree has gained particular interest for silvopasture systems due to the production of edible, high-sugar seedpods that livestock may consume after pod drop. Two studies were conducted within an active sheep and honeylocust silvopasture to (1) estimate nutritional variability of seedpods among MW trees, and to (2) determine seedpod yields and seasonal production variation of MW trees. Seedpods were harvested from each pod-bearing MW in October 2008 and 2009, just prior to pod drop. Nutritional characteristics such as detergent fibers, crude protein, in vitro digestibility, and sugar concentrations were determined for fractionated husk and seed components. Further, MW fodder yields were estimated in 2008, 2009, and 2010 through field measurements and tree yield classification. Both ground husks and seeds were low in fiber (273 and 132 g kg −1 neutral detergent fiber, respectively) and highly digestible (787 and 963 g kg −1 , respectively). Seed husk sugar concentrations averaged 223 g kg −1 . Based on the nutritive fractions assayed, whole ground MW seedpods grown in Virginia have a nutritional profile comparable to that of ground whole-ear dent corn ( Zea mays L.) or oat ( Avena sativa L.) grain. Nearly all MW trees displayed some form of alternate-bearing pattern. Average dry matter yields of pod-bearing trees were 15.8, 4.8, and 14.7 kg tree −1 in 2008, 2009, and 2010, respectively. Pod yield and quality indicate MW honeylocust trees have good potential as fodder-bearing trees in temperate silvopasture systems.