Summary
The shallow marine subtropical Northern Bay of Safaga is composed of a complex pattern of sedimentary facies that are generally rich in molluscs. Thirteen divertaken bulk-samples from various sites (reef slopes, sand between coral patches, muddy sand, mud, sandy seagrass, muddy seagrass, mangrove channel) at water depths ranging from shallow subtidal to 40m were investigated with regard to their mollusc fauna >1mm, which was separated into fragments and whole individuals.
Fragments make up more than 88% of the total mollusc remains of the samples, and their proportions correspond to characteristics of the sedimentary facies. The whole individuals were differentiated into 622 taxa. The most common taxon,Rissoina cerithiiformis, represented more than 5% of the total mollusc content in the samples. The main part of the fauna consists of micromolluscs, including both small adults and juveniles. Based on the results of cluster-, correspondence-, and factor analyses the fauna was grouped into several associations, each characterizing a sedimentary facies: (1) “Rhinoclavis sordidula—Corbula erythraeensis-Pseudominolia nedyma association” characterizes mud. (2) “Microcirce sp.—Leptomyaria sp. association” characterizes muddy sand. (3)”Smaragdia spp.-Perrinia stellata—Anachis exilis—assemblage” characterizes sandy seagrass. (4) “Crenella striatissima—Rastafaria calypso—Cardiates-assemblage” characterizes muddy seagrass. (5) “Glycymeris spp.-Parvicardium sueziensis-Diala spp.-assemblage” characterizes sand between coral patches. (6) “Rissoina spp.-Triphoridae —Ostreoidea-assemblage” characterizes reef slopes. (7) “Potamides conicus—Siphonaria sp. 2—assemblage” characterizes the mangrove.
The seagrass fauna is related to those of sand between coral patches and reef slopes with respect to gastropod assemblages, numbers of taxa and diversity indices, and to the muddy sand fauna on the basis of bivalve assemblages and feeding strategies of bivalves. The mangrove assemblage is related to those of sand between coral patches and the reef slope with respect to taxonomic composition and feeding strategies of bivalves, but has a strong relationship to those of the fine-grained sediments when considering diversity indices. Reef slope assemblages are closely related to that of sand between coral patches in all respects, except life habits of bivalves, which distincly separates the reef slope facies from all others.
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References
Antia, D.D.J. (1977): A comparison of diversity and trophic nuclei of live and dead molluscan faunas from the Essex Chenier plain, England.—Paleobiology3, 404–414, 3 Fig., 3 Tabs., Chicago
Aller, R.C. (1982): Carbonate dissolution in nearshore terrigenous muds: the role of physical and biological reworking.—Journal of Geology90, 79–95, 11 Figs., Chicago
Arnoud, P.M. &Thomassin, B.A. (1990): Habits and morphological adaptations of mytilids (Mollusca: Bivalvia) from coastal and reefal environments in south-west Malagasia (Indian Ocean). In:Brian Morton (ed.): The Bivalvia-Proceedings of a Memorial Symposium in Honour of Sir Charles Maurice Yonge, Edinburgh, 1986.—333–344, 4 Figs. Hong Kong (Hong Kong Univ. Press)
Austin, A.D., Austin, S.A. andSale, P.F. (1980): Community structure of the fauna associated with the coralPocillopora damicornis (L.) on the Great Barrier Reef. Aust. J. Mar. Freshwater Res.31, 163–174, Melbourne
Ayal, Y. &Safriel, U.N. (1981): Species composition, geographical distribution and habitat characteristics of rocky intertidal Cerithiidae (Gastropoda: Prosobranchia) along the Red Sea shores of Sinai. Argamon7(5), 53–72, 2 Figs., 3 Tabs., Haifa
Bandel, K. (1991a): Character of a microgastropod fauna from a carbonate sand of Cebu (Philippines).—Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg71, 441–485 8 Pls., 36 Figs., Hamburg
— (1991b): Schlitzbandschnecken mit perlmutteriger Schale aus den triassischen St. Cassian-Schichten der Dolomiten.— Ann. Naturhist. Mus. Wien92(A), 1–53, 17 Pls., 10 Figs, Wien
— (1993): Trochomorpha aus der triassischen St. Cassian-Formation (Gastropoda, Dolomiten). Ann. Naturhist. Mus. Wien95(A), 1–99 16 Pls., Wien
Benzecri, P.J. (1973): L'analyse de correspondances. L'Analyse de Données, 2, Paris (Dunod)
Bosch, D.T., Dance, S.P., Moolenbeek, R.G. &Oliver, P.G. (1995): Seashells of Eastern Arabia.—296pp. Dubai (Motivate Publishing).
Bratcher, T. &Cernohorsky, W.O. (1987): Living Terebras of the World. A Monograph of the Recent Terebridae of the World. Melbourne (Florida), Burlington (Mass.). American Malacologists.
Cadèe, G.C. (1994): Eider, shelduck, and other predators, the main producers of shell fragments in the Wadden Sea.— Palaeontology37/1, 181–202, 1 Pl., 13 Figs. 3 Tabs., London
Calef, C.E. &Hancock, N.J. (1974): Wenlock and Lludlow marine communities in Wales and the Welsh borderland. —Palaeontology17:779–810, 1 Pl., 7 Figs., 13 Tabs., London
Carthew, R. &Bosence, D. (1986): Community preservation in recent shell-gravels. English Channel.—Palaeontology29(2), 243–268, 7 Figs., 4 Tabs., 2 Apps., London
Cernohorsky, W.O. (1984): Systematics of the Family Nassariidae (Mollusca: Gastropoda). Bull. Auckland Inst. Mus.14, 51 Pls., 173 Figs., Auckland
Crame, J.A. (1986): Late Pleistocene molluscan assemblages from the coral reefs of the Kenya coast.—Coral reefs4, 183–196, 6 Figs., 2 Tabs., Berlin
Cummins, H., Powell, E.N., Standon, R.J. Jr. &Staff, G. (1986): The size-frequency distribution in Palaeoecology: effects of taphonomic processes during formation of molluscan death assemblages in Texas bays.—Palaeontology29(3), 495–518: 12 Figs., 7 Tab., London
Davies, D.J., Powell, E.N., Stanton, R.J. Jr. (1989a): Taphonomic signature as a function of environmental process: shells and shell beds in a hurricane-influenced inlet on the Texas coast.-Palaeogeography, Palaeoclimatology, Palaeoecology72, 317–356, Fig. 11. Tab. 26, Amsterdam
Davies, D.J., Powell, E.N., Stanton, R.J. Jr. (1989b): Relative rates of shell dissolution and net sediment accumulation-a commentary: can shell beds form by the gradual accumulation of biogenic debris on the sea floor?.—Lethaia22 207–212, 1 Fig., Oslo
Davies, D.J., Staff, G.M., Callender, W.R., Powell, E.N. (1990): Description of a Quantitative Approach to Taphonomy and Taphofacies Analysis: All Dead Things Are Not Created Equal.—In:Miller, W. (ed.): Paleocommunity temporal dynamics: The long-term development of multispecies assemblies— The Paleontological Society Special Publication5, 328–350, 5 Pls., 1 Fig., 1 Tab., Knoxville (The University of Tennessee)
Dullo, W.Chr. (1983): Diagenesis of fossils of the miocene Leitha Limestone of the Paratethys, Austria: an example for faunal modifications due to the changing diagnetic environment.—Facies8, 1–112, 15 Pl., 22 Fig., 2 Tab., Erlangen
— (1984): Progressive diagnetic sequence of aragonite structures: Pleistocene coral reefs and their modern counterparts on the eastern Red Sea coast, Saudi Arabia.—Palaeontographica Americana54, 254–260, 5 Figs., Ithaca
— (1990): Facies, fossil record, and age of Pleistocene reefs from the Red Sea.—Facies22, 1–46, 13 Pl., 21 Figs., 1 Tab., Erlangen
Dworschak, P.C. &Pervesler, P. (1988): Burrows ofCallianassa bouvieri NOBILI 1904 from Safaga (Egypt, Red Sea) with some remarks on the Biology of the species.—Senckenbergiana marit.20(1/2): 1–17, 2 Pls., 5 Figs., 1 Tabs., Frankfurt
Ekdale, A.A. (1977): Quantitative Paleoecological Aspects of Modern Marine Mollusk Distribution, Northeast Yucatán Coast, Mexico.—In:Frost, S.H., Weiss, M.P. & Saunders, J.P. (eds.): Reefs and related carbonates—ecology and sedimentology.-Studies in Geology4, 195–207, 12 Figs Tulsa
Flessa, K.W. (1993): Time-averaging and temporal resolution in Recent marine shelly faunas.—In:Kidwell, S.M. &Behrensmeyer, A.K. (eds.): Taphonomic approaches to time resolution in fossil assemblages.—The Paleontological Society Special Publication6 9–33, 3 Figs., 1 Tab., Knoxville (The University of Tennessee)
Flessa K.W. &Kowalewski M. (1994): Shell survival and time-averaging in nearshore and shelf environments: estimates from the radiocarbon literature.—Lethaia27, 153–165, 4 Figs., 3 Tabs., Oslo
Flessa, K.W., Cutler, A.H. &Meldahl, K.H. (1993): Time and taphonomy: quantitative estimates of time-averaging and stratigraphic disorder in a shallow marine habitat.—Paleobiology19(2): 266–286, 7 Figs., 3 Tabs., Chicago
Flessa, K.W., Kowalewski, M. &Walker, S.E. (1992): Postcollection taphonomy: shell destruction and the chevrolet.— Palaios7, 553–554, 1 Tab., Tulsa
Frank, P.W. (1969): Growth rates and longevity of some gastropod molluscs on the coral reef at Heron Island. Oecologia2, 232–250, 3 Figs., 3 Tabs., Berlin
Fretter, V. (1975):Umbonium vestiarium, a filter-feeding trochid. Journal of Zoology177, 541–552, 4 Figs., London
Fürsich, F.T. (1977): Corallian (Upper Jurassic) marine benthic associations from England and Normandy).—Palaeontology20, 337–386, London
— (1978): The influence of faunal condensation and mixing on the preservation of fossil benthic communities.—Lethaia11, 243–250, 7 Figs., Oslo
Fürsich, F.T. & Aberhan, M. (1990): Significance of time-averaging for palaeocommunity analysis.—Lethaia23, 143–152, 6 Figs., Oslo
Gabriel, K.R. (1971): The biplot—graphic display of matrices with application to principal component analsis. Biometrika58, 435–467, London
Glinsky, N.L. &Bennington, J.B. (1994): Estimating numbers of whole individuals from collections of body parts: a taphonomic limitation of the paleontological record.— Paleobiology20(2), 245–258, 5 Figs., 1 Tab., Chicago
Glover, E.A. &Taylor, J.D. (1997): New species and records ofRastafaria andMegaxinus (Bivalvia: Lucinidae) from the Western Indian Ocean and Red Sea, with reappraisal ofMegaxinus.—Journal of Conchology36(1), 1–18, 7 Figs., London
Hadfield, M.G. (1976): Molluscs associated with living corals.— Micronesica12, 133–148, 1 Tab., Agaña (Guam)
Haunold, T.G., Baal, C. &Piller, W.E. (1997): Benthic Foraminiferan Associations in the Northern Bay of Safaga, Red Sea, Egypt.—Marine Micropaleontology29, 185–210, 16 Figs., 2 Tabs., New York
Hickman, C.S. (1985): Comparative morphology and ecology of free-living suspension-feeding gastropods from Hong Kong.— In:Morton, B. &Dudgeon, D. (eds.): Proceedings of the Second International Workshop on the Malacofauna of Hong Kong and Southern China, Hong Kong, 1983,—217–234, 1 Pl., 4 Figs., Hong Kong, (Univ. Press)
Hill, M.O. (1973): Reciprocal averaging: an eigenvector method of ordination. Journal of Ecology,61, 237–249, 2 Figs., Cambridge
Holtzmann, R.C. (1979): Maximum likelihood estimation of fossil assemblage composition. Paleobiology5(2), 77–89, 5 Figs., 1 Tab., Chicago
Houbrick, R.S. (1978): The family Cerithiidae in the Indo-Pacific. Part 1: The generaRhinoclavis, Pseudovertagus andClavocerithium.—Monographs of Marine Mollusca, 1:1–130, 98 Pls., American Malacologists, Inc: Greenville, Delaware
Houbrick, R. S. (1992): Monograph of the GenusCerithium Bruguière in the Indo-Pacific (Cerithiidae: Prosobranchia).—Smithsonian Contribution to Zoology510, 211pp. 145 Figs. 45 Tabs., Washington
Janssen, R. (1981): Mollusken-Assoziationen und Biotope im norddeutschen Oberoligozän.—Natur und Museum111, 70–78, 8 Figs., Frankfurt a. M.
Joreskog, K.G., Klovan, J.E. andReyment, R.A. (1976): Geological Factor Analysis.—XII+ 178 pp, Amsterdam (Elsevier)
Kay, E.A. (1979): Hawaiian marine shells. Reef and shore fauna of Hawaii, section 4: Mollusca.—653 pp. 195 Figs., Honolulu
Kay, E.A. &Switzer, M.F. (1974): Molluscan distribution patterns in Fanning Island Lagoon and a comparison of the mollusks of the lagoon and the seaward reefs.—Pac. Sci.28(2), 275–295, 6 Figs., 5 Tabs., Hawaii
Kennard, M.C. &Smith, A.J. (1961): A simple micro-sample splitter.—Journal of Paleontology35, 396–397, 2 Figs. 1 Tab., Tulsa
Kidwell, S.M. (1993): Patterns of time-averaging in the shallow marine fossil record.—In: Kidwell, S.M. & Behrensmeyer, A.K. (eds.): Taphonomic approaches to time resolution in fossil assemblages.—The Paleontological Society Special Publication6, 275–300, 6 Figs., 1 Tab., Knoxville (The University of Tennessee)
Kidwell, S.M. &Bosence, D.W.J. (1991): Taphonomy and time-averaging of marine shelly faunas. In:Allison, P.A. &Briggs, D.E.G. (eds.): Taphonomy. Releasing the data locked in the fossil record.—115–209, 10 Figs., 9 Tabs., New York (Plenum)
Kidwell, S.M. &Flessa, K.W. (1995): The quality of the fossil record: Populations, species, and communities.—Annu. Rev. Ecol. Syst.26, 269–299, 2 Figs., Palo Alto
Kidwell, S.M., Fürsich, F.T. &Aigner, T. (1986): Conceptual framework for the analysis of fossil concentrations.—Palaios1, 228–238, 5 Figs., 1 Tab., Tulsa
Kleemann, K. (1990): Coral Associations, Biocorrosion, and Space Competition inPedum spondyloideum (Gmelin) (Pectinacea, Bivalvia): Publs. Staz. zool. Napolil: Marine Ecology11, 77–94, 19 Figs., 1 Tab., Berlin
Kleemann, K. (1992): Coral Communities and Coral-Bivalve Associations in the Northern Red Sea at Safaga. Egypt: Facies26, 1–10, 3 Pl., 1 Fig., 1 Tab., Erlangen
Kleemann, K. (1995): Associations of coral and coral boring bivalves: Lizard Island (Great Barrier Reef, Australia) versus Safaga (N Red Sea).—Beiträge zur Paläontologie20, 31–39, 1 Pl., 2 Figs., 1 Tab., 1 App., Wien
Kohn, A.J. (1983): Feeding Biology of Gastropods.—In:Saleuddin, A.S.M., andWilbur, K.M. (eds.) The Mollusca, Vol. 5 Physiology, Part 2.—1–63, 2 Figs., 2 Tabs. New York (Academic Press)
Kowalewski, M. (1996): Time-averaging, overcompleteness, and the geological record.—The Journal of Geology104:317–326, 3 Figs., Chicago
— (1997): The reciprocal taphonomic model.—Lethaia30, 86–88, 1 Tab., Oslo
Krebs, Ch.J. (1989): Ecological Methodology.—X+654 pp., New York (Harper & Row)
Krzanowski, W.J. &F.H.C. Marriott (1995): Multivariate Analysis, Part 2. Kendall's Library of Statistics 2. VIII+280 pp. 17 Figs., London (Arnold)
Lee, S.Y., Morton, B. (1985): The Hong Kong Mytilidae. In:Morton, B. &Dudgeon, D. (eds.): Proceedings of the Second International Workshop on the Malacofauna of Hong Kong and Southern China, Hong Kong, 1983.—49–76, 3 Figs., 5 Pls., 3 Tabs., Hong Kong, (Hong Kong Univ. Press)
Le Pennec, M., Beninger, P.G. &Herry, A. (1995): Feeding and digestive adaptations of bivalve molluscs to sulphide-rich habitats.—Comparative Biochemistry and Physiology111A: 183–189, London.
Linder, A. &Berchtold, W. (1976): Statistische Auswertung von Prozentzahlen.—232 pp. Basel (Birkhäuser)
Magurran, A.E. (1988): Ecological Diversity and Its Measurement.—X+179 pp., London-Sydney (Croom Helm)
Mansour, A.M., Piller, W.E. &Lengauer, C.L. (1995): Quantitative mineralogical analyses of carbonate rich sediments by X-ray powder diffraction.—Powder Diffraction10 (2): 112–116, 2 Figs., 4 Tabs., Woodbury
Marshall, B.A. (1983): A Revision of the Recent Triphoridae of Southern Australia (Mollusca: Gastropoda)—Records of the Australian Museum, Supplement2, 119 pp., 33 Figs. 44 Tabs., 1 App., Sydney
Mastaller, M. (1978): The marine mollusean assamblages of Port Sudan. Red Sea. Zoologische Mededelingen53, 117–144, 1 Fig., 1 Tab., Leiden
Mastaller, M. (1988): Late Miocene deep-water Mollusca from the Still water Mudstone at Graymouth, Westland, New Zealand: Paleoecology and systematics.—New Zealand Geological Survey Paleontological Bulletin55, 120 pp., 13 Pls., 3 Figs., 11 Tabs, Lower Hutt
Maxwell, P.A. (1992): Eocene mollusca from the vicinity of McCulloch's Bridge, Waihao River, South Canterbury, New Zealand: Paleoecology and systematics.—New Zealand Geological Survey Paleontological Bulletin65. 280 pp, Lower Hutt
McCall, P.L. &Tevesz, M.J.S. (1983): Soft-bottom succession and the fossil record.—In:Tevesz, M.J.S. &McCall, P.L. (eds.): Biotic interactions in recent and fossil benthic communities.—157–196 New York (Plenum Press)
Meldahl, K.H. (1987): Sedimentologic and taphonomic implications of biogenic stratification.—Palaios,2, 350–358, 7 Figs., Tulsa
Meldahl, K.H. &Flessa, K.W. (1990): Taphonomic pathways and comparative biofacies and taphofacies in a recent intertidal/shallow shelf environment.—Lethaia23, 43–60, 11 Figs., 1 Tab., Olso
Meldahl, K.H., Flessa, K.W. &Cutler, A.H. (1997): Timeaveraging and postmortem skeletal survival in benthic fossil assemblages: quantitative comparisons among Holocene environments.—Paleobiology23(2): 207–229, 7 Figs., 3 Tabs., Chicago
Morton, B. (1983): Feeding and digestion in Bivalvia. In:Saleuddin, A.S.M. &Wilbur, K.M. (eds.): The Moliusea, Vol. 5 Physiology, Part 2.—65–147, 22 Figs. New York (Academic Press)
Myrick, J.L. &Flessa, K.W. (1996): Bioturbation rates in Bahía La Choya, Sonora, Mexico.—Ciencias Marinas,22(1): 23–46, 10 Figs., 2 Tabs., Mexico
Nebelsick, J.H. (1992a): The Northern Bay of Safaga (Red Sea, Egypt): an actuopalaeontological approach. III. Distribution of echinoids. Beiträge zur Paläontologie von Österreich17, 5–79, 8 Pls., 37 Figs., 6 Tables, Wien
— (1992b): Echinoid Distribution by Fragment Identification in the Northern Bay of Safaga, Red Sea, Egypt.—Palaios7(2): 316–328, 8 Figs., 2 Tabs., Tulsa
— (1995a): Comparative taphonomy of Clypeasteroids.—Eclogae geol. Helv.88/3: 685–693, 7 Figs., Basel
— (1995b): Uses and limitations of actuopalaeontological investigations on echinoids.—Geobios18, 329–336, 7. Figs., Lyon
— (1996): Biodiversity of shallow-water Red Sea echinoids: Implications for the fossil record.—J. mar. biol. Ass. U.K.76, 185–194, 4 Figs., 1 Tab., London
Nebelsick, J.H. &Kampfer, S. (1994): Taphonomy of Clypeaster humilis and Echinodiscus auritus (Echinoidea, Clypeasteroidea) from the Red Sea.—In:David, B., Guille, A., Feral, J.-P. &Roux, M. (eds.): Echinoderms through time, 803–808, 1 Pl., 4 Figs. Rotterdam (Balkema)
Oliver, P.G. (1992): Bivalved Seashells of the Red Sea—330 pp., Wiesbaden (Hemmen)
Orloci, L. (1967): Geometric models in ecology. The theory and application of some ordination methods.—Journal of Ecology54, 193–215, 15 Figs., 2 Tabs., Cambridge
Perron, F.E. (1975): Carnivorous Calliostoma (Prosobranchia: Trochidae) from the northeastern Pacific.—Veliger18, 52–54, 1 Pl., Berkeley
Piller, W.E. (1994): The Northern Bay of Safaga (Red Sea, Egypt): an actuopalaeonotological approach. IV. Thin section analysis. Beiträge zur Paläontologie18, 1–73, 18 Pls., 19 Figs., 5 Tables, Wien
Piller, W.E. &Mansour, A. (1990): The Northern Bay of Safaga (Red Sea, Egypt): an actuopalaeontological approach. II. Sediment analyses and sedimentary facies.—Beiträge zur Paläontologie von Österreich16, 1–102, 55 Figs., 19 Tabs., Wien
Piller, W.E. &Massour, A. (1994): Origin and transport of noncarbonate sediments in a carbonate-dominated environment (Northern Safaga Bay, Egypt).—Abhandlungen der Geologischen Bundesanstalt50, 369–379, 1 Pl., 6 Figs., 3 Tabs., Wien.
Piller, W.E. &Pervesler, P. (1989): The Northern Bay of Safaga (Red Sea, Egypt): An Actuopalaeontological Approach. I. Topography and Bottom Facies.—Beiträge zur Paläontologie von Österreich15, 103–147, 10 Pls., 8 Figs., 1 Table, Wien
Piller, W.E. &Rasser, M. (1996): Rhodolith formation induced by reef erosion in the Red Sea, Egypt.—Coral Reefs15, 191–198, 6 Figs., Berlin
Pielou, E.C. (1969): An Introduction to Mathematical Ecology. VIII+286 pp., New York (Wiley)
Ponder, W. F. (1985): A review of the genera of the Rissoidae (Mollusca: Mesogastropoda: Rissoacea).—Rec. Austr. Mus. Suppl.4, 1–221 153 Figs., Sydney.
Powell, E.N. &Davies, D.J. (1990): When is an “old” shell really “old”?.—Journal of Geology98, 823–844 15 Figs., Chicago
Powell, E.N., Staff, G.M., Davies, D.J. &Callender, W.R. (1989): Macrobenthic Death Assemblages in Modern Marine Environments: Formation, Interpretation, and Application.—Aquatic sciences1/4, 555–589, 2 Figs., 8 Tabs., London
Powell, E.N. &Stantos, R.J., Jr. (1985): Estimating biomass and energy flow of molluses in palaeo-communities.—Palaeontology28, 1–34, 14 Figs., 6 Tabs., London
— & — (1995): The application of guild and tier structure and energy flow in paleoecologic analysis: an example using parautochthonous death assemblages from a variable salinity bay.—Historical Biology10, 281–327, 12 Fig., 5 Tab., Amsterdam
— & — (1996): The application of siz-frequency distribution and energy flow in paleoecological analysis: An example using parautochthonous death assemblages from a variable salinity bay.—Palaeogeography, Palaeoclimatology, Palaeoecology124, 195–231, 26 Figs., 3 Tabs., Amsterdam.
Powell, E.N., Stanton, R.J., Jr., Logan, A. &Craig, M.A. (1992): Preservation of Mollusca in Copano Bay, Texas. The long-term record.—Palaeogeography, Palaeoclimatology, Palaeoecology95, 209–228, 20 Figs., Amsterdam
Rasser, M. &Piller, W.E. (1997): Depth distribution of calcareous encrusting associations in the northern Red Sea (Safaga, Egypt) and their geological implications.—Proceedings of the 8th International Coral Reef Symposium 1, 743–748, 9 Figs., Panama
Riegl, B. &Piller, W.E. (1997): Distribution of coral associations in Northern Safaga Bay (Red Sea, Egypt).—Facies36, 141–162, 8 Pls., 3 Figs., 1 Tab., Erlangen
Runnegar, B. (1996): Early evolution of the mollusca: the fossil record.—In:Taylor, J. D. (ed.): Origin and evolutionary radiation of the mollusca.—77–87, 5 Figs. Oxford (Oxford Univ. Press)
Salazar-Jimenez, A., Frey, R.W. &Howard, J.D. (1982): Concavity orientations of bivalve shells in estuarine and nearshore shelf sediments, Georgia.—J. Sediment. Petrol.52, 565–586, 12 Figs., 2 Tabs., Tulsa
Safriel, U.N., Felsenburg, T. &Gilboa, A. (1980): The distribution ofBrachidontes variabilis (Krauss) along the Red Sea coasts of Sinai.—Argamon7/3, 31–43, 5 Tabs., Haifa
Scarabino, V. (1995): Scaphopoda of the tropical Pacific and Indian Oceans, with description of 3 new genera and 42 new species.—In:Bouchet, P., (ed.): Résultats des Campagnes MUSORSTOM, Vol. 14.—Mém. Mus.natn. Hist. nat.167, 189–397, 175 Figs., 2 Tabs., Pars
Schuhmacher, H. (1993): Impact of Some Corallivorous snails on Stony Corals in the Red Sea Proceedings of the Seventh International Coral Reef Symposium, Guam 1992,2, 840–846, 3 Figs., 2 Tabs., Guam
Sharabati, D. (1984): Red Sea Shells.—127 pp, 49 Pls., London (Routledge & Keegan)
Sheppard, A.L.S. (1984): The molluscan fauna of Chagos (Indian Ocean) and an analysis of its broad distribution patterns.—Coral Reefs3, 43–50, 3 Figs., 1 Tab., 1 App., Berlin
Staff, G.M. &Powell, E.N. (1988): The paleoecological significance of diversity: the effect of time averaging and differential preservation on macroinvertebrate species richness in death assemblages.—Palaeogeography, Palaeoclimatology, Palaeoecology63, 73–89, 11 Figs., 2 Tabs., Amsterdam
Staff, G.M., Stanton, R.J. Jr., Powell, E.N. &Cummins, H. (1986): Time-averaging, taphomomy, and their impact on paleocommunity reconstruction: Death assemblages in Texas bays.—Geological Society of America Bulletin97, 428–433, 14 Figs., 2 Tabs., Boulder
Stanton, R.J., Jr., Powell, E.N. &Nelson, P.C. (1981): The role of carnivorous gastropods in the trophic analysis of a fossil community.—Malacologia,20/2, 451–469, 14 Figs., 2 Tabs., Ann Arbor
Taylor, J.D. (1968): Coral reef and associated invertebrate communities (mainly molluscan) around Mahé, Seychelles.—Phil. Trans. R. Soc. London254 (Ser. B), 129–206, 20 Figs., London
— (1971): Intertidal zonation of Aldabra Atoll.—Philosophical Transactions of the Royal Society,260 (Ser. B), 173–213, London
— (1976): Habitats, abundance and diets of Muricacean gastropods at Aldabra Atoll.—Zoological Journal of the Linnean Society59, 155–193, 16 Figs., 12 Tabs., London
Taylor, J.D., &Reid, D.G. (1984): The abundance and trophic classification of molluscs upon coral reefs in the Sudanese Red Sea: Journal of Natural History18, 175–209, 11 Figs., 7 Tabs., 1 App., London
Trewin N.H. &Welsh W. (1976): Formation and composition of a graded estuarine shell bed.—Palaeogeogr., Palaeoclimatol., Palaeoecol.19, 219–230, 5 Figs., Amsterdam
Valentine, J.W. (1989): How god was the fossil record? Clues from the Californian Pleistocene.—Paleobiology15/2, 83–94, 1 Fig., 1 Tab., Chicago
Vermeli, G.J. (1983): Traces and trends of predation, with special reference to bivalved animals.—Palaeontology26, 455–465, 1 Fig., London
Walker, K.R. (1972): Trophic analysis: a method for studying the function of ancient communities.—J. Paleontol.46, 82–93, 11 Tabs., Tulsa
Ward, J.H. (1969): Hierarchical grouping to optimize an objective function.—J. Amer. Stat. Assoc.58, 236–244, 3 Figs., Washington, D.C.
Warme, J.E. (1969) Live and dead molluscs in a coastal lagoon.—Journal of Paleontology43/1, 141–150, 2 Figs., 2 Tabs., Tulsa
Westhoff, V. &Vander Maarel, E. (1973): The Braun-Blanquet approach.—In:R.H. Whittaker (ed.): Handbook of vegetation science 5, Ordination and Classification of Communities.—619–726, 6 Figs., 7 Tabs., The Hague (Junk)
Zuschin, M. & Oliver, P.G. (submitted): Additions to the bivalve fauna of the Red Sea with descriptions of new species of Limopsidae, Limidae, Semelidae and Veneridae.—Journal of conchology, London
Zuschin, M. &Piller, W.E. (1997a): Gastropod shells recycled—an example from a rocky tidal flat in the Northern Bay of Safaga (Red Sea, Egypt).—Lethaia30, 127–134, 6 Figs., 4 Tabs., Oslo
— & — (1997b): Bivalve distribution on coral carpets in the northern Bay of Safaga (Red Sea, Egypt) and its relation to environmental parameter.—Facies37, 183–194, 2 Pls., 5 Figs., 6 Tabs., Erlangen
— & — (1997c): Molluscan hard-substrate associations in the northern Red Sea.—Pubbl. Staz zool. Napoli: Marine Ecology,18/4, 361–378, 3 Figs., 3 Tabs. Berlin
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Zuschin, M., Hohenegger, J. Subtropical coral-reef associated sedimentary facies characterized by molluscs (Northern Bay of Safaga, Red Sea, Egypt). Facies 38, 229–254 (1998). https://doi.org/10.1007/BF02537367
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DOI: https://doi.org/10.1007/BF02537367